---
title: "Nature 2021 — all accepted papers with summaries and citations"
description: "All 3,641 papers of Nature 2021 · page 1 of 37, the most cited first; those with our summary link to it."
canonical: "https://research.elamur.ai/free/en/venues/nature/2021/papers/"
---
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# Nature 2021: all accepted papers

3,641 papers · page 1 of 37 · the most cited first · [← Nature 2021](https://md.research.elamur.ai/free/en/venues/nature/2021/)

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1. **[Highly accurate protein structure prediction with AlphaFold](https://md.research.elamur.ai/free/en/papers/jumper-2021-highly-accurate-protein-structure-prediction/)**John Jumper, Richard Evans, Alexander Pritzel et al.*43,269 citations · our summary*
2. **[Perovskite solar cells with atomically coherent interlayers on SnO2 electrodes ↗](https://doi.org/10.1038/s41586-021-03964-8)**Hanul Min, Do Yoon Lee, Junu Kim et al.*3,174 citations*
3. **[Towards complete and error-free genome assemblies of all vertebrate species ↗](https://doi.org/10.1038/s41586-021-03451-0)**Arang Rhie, Shane McCarthy, Olivier D. Fedrigo et al.*3,098 citations*
4. **[Pseudo-halide anion engineering for α-FAPbI3 perovskite solar cells ↗](https://doi.org/10.1038/s41586-021-03406-5)**Jaeki Jeong, Minjin Kim, Jongdeuk Seo et al.*3,072 citations*
5. **[Highly accurate protein structure prediction for the human proteome ↗](https://doi.org/10.1038/s41586-021-03828-1)**Kathryn Tunyasuvunakool, Jonas Adler, Zachary Wu et al.*2,998 citations*
6. **[Efficient perovskite solar cells via improved carrier management ↗](https://doi.org/10.1038/s41586-021-03285-w)**Jason J. Yoo, Gabkyung Seo, Matthew R. Chua et al.*2,931 citations*
7. **[Designing the next generation of proton-exchange membrane fuel cells ↗](https://doi.org/10.1038/s41586-021-03482-7)**Kui Jiao, Jin Xuan, Qing Du et al.*2,915 citations*
8. **[The changing landscape of atherosclerosis ↗](https://doi.org/10.1038/s41586-021-03392-8)**Peter Libby*2,838 citations*
9. **[Detection of a SARS-CoV-2 variant of concern in South Africa ↗](https://doi.org/10.1038/s41586-021-03402-9)**Houriiyah Tegally, Eduan Wilkinson, Marta Giovanetti et al.*2,509 citations*
10. **[Antibody resistance of SARS-CoV-2 variants B.1.351 and B.1.1.7 ↗](https://doi.org/10.1038/s41586-021-03398-2)**Pengfei Wang, Manoj S. Nair, Lihong Liu et al.*2,310 citations*
11. **[A 20-year retrospective review of global aquaculture ↗](https://doi.org/10.1038/s41586-021-03308-6)**Rosamond L. Naylor, Ronald W. Hardy, Alejandro H. Buschmann et al.*2,233 citations*
12. **[Sequencing of 53,831 diverse genomes from the NHLBI TOPMed Program ↗](https://doi.org/10.1038/s41586-021-03205-y)**Daniel Taliun, Daniel Harris, Michael D. Kessler et al.*2,192 citations*
13. **[Omicron escapes the majority of existing SARS-CoV-2 neutralizing antibodies ↗](https://doi.org/10.1038/s41586-021-04385-3)**Yunlong Cao, Jing Wang, Fanchong Jian et al.*1,996 citations*
14. **[Reduced sensitivity of SARS-CoV-2 variant Delta to antibody neutralization ↗](https://doi.org/10.1038/s41586-021-03777-9)**Delphine Planas, David L. Veyer, Artem Baidaliuk et al.*1,987 citations*
15. **[DHODH-mediated ferroptosis defence is a targetable vulnerability in cancer ↗](https://doi.org/10.1038/s41586-021-03539-7)**Chao Mao, Xiaoguang Liu, Yilei Zhang et al.*1,900 citations*
16. **[Accelerated global glacier mass loss in the early twenty-first century ↗](https://doi.org/10.1038/s41586-021-03436-z)**Romain Hugonnet, Robert McNabb, Étienne Berthier et al.*1,889 citations*
17. **[Parallel convolutional processing using an integrated photonic tensor core ↗](https://doi.org/10.1038/s41586-020-03070-1)**J. Feldmann, N. Youngblood, M. Karpov et al.*1,874 citations*
18. **[Photocatalytic solar hydrogen production from water on a 100-m2 scale ↗](https://doi.org/10.1038/s41586-021-03907-3)**Hiroshi Nishiyama, Taro Yamada, Mamiko Nakabayashi et al.*1,776 citations*
19. **[In situ Raman spectroscopy reveals the structure and dissociation of interfacial water ↗](https://doi.org/10.1038/s41586-021-04068-z)**Yao-Hui Wang, Shisheng Zheng, Weimin Yang et al.*1,703 citations*
20. **[Exploring tissue architecture using spatial transcriptomics ↗](https://doi.org/10.1038/s41586-021-03634-9)**Anjali N. Rao, Dalia Barkley, Gustavo S. França et al.*1,669 citations*
21. **[Evolution of antibody immunity to SARS-CoV-2 ↗](https://doi.org/10.1038/s41586-021-03207-w)**Christian Gaebler, Zijun Wang, Julio C. C. Lorenzi et al.*1,665 citations*
22. **[Developing fibrillated cellulose as a sustainable technological material ↗](https://doi.org/10.1038/s41586-020-03167-7)**Tian Li, Chaoji Chen, Alexandra H. Brozena et al.*1,663 citations*
23. **[Assessing transmissibility of SARS-CoV-2 lineage B.1.1.7 in England ↗](https://doi.org/10.1038/s41586-021-03470-x)**Erik Volz, Swapnil Mishra, Meera A Chand et al.*1,601 citations*
24. **[Strong tough hydrogels via the synergy of freeze-casting and salting out ↗](https://doi.org/10.1038/s41586-021-03212-z)**Mutian Hua, Shuwang Wu, Yanfei Ma et al.*1,519 citations*
25. **[11 TOPS photonic convolutional accelerator for optical neural networks ↗](https://doi.org/10.1038/s41586-020-03063-0)**Xingyuan Xu, Mengxi Tan, Bill Corcoran et al.*1,518 citations*
26. **[Considerable escape of SARS-CoV-2 Omicron to antibody neutralization ↗](https://doi.org/10.1038/s41586-021-04389-z)**Delphine Planas, Nell Saunders, Piet Maes et al.*1,511 citations*
27. **[High-dimensional characterization of post-acute sequelae of COVID-19 ↗](https://doi.org/10.1038/s41586-021-03553-9)**Ziyad Al‐Aly, Yan Xie, Benjamin C. Bowe*1,511 citations*
28. **[Satellite imaging reveals increased proportion of population exposed to floods ↗](https://doi.org/10.1038/s41586-021-03695-w)**Beth Tellman, Jonathan A. Sullivan, C Kuhn et al.*1,463 citations*
29. **[Ultralow contact resistance between semimetal and monolayer semiconductors ↗](https://doi.org/10.1038/s41586-021-03472-9)**Pin‐Chun Shen, Cong Su, Yuxuan Cosmi Lin et al.*1,402 citations*
30. **[High and rising economic costs of biological invasions worldwide ↗](https://doi.org/10.1038/s41586-021-03405-6)**Christophe Amidi Diagne, Boris Leroy, Anne‐Charlotte Vaissière et al.*1,398 citations*
31. **[Promises and prospects of two-dimensional transistors ↗](https://doi.org/10.1038/s41586-021-03339-z)**Yuan Liu, Xidong Duan, Xidong Duan et al.*1,356 citations*
32. **[Small-molecule inhibition of METTL3 as a strategy against myeloid leukaemia ↗](https://doi.org/10.1038/s41586-021-03536-w)**Eliza Yankova, Wesley Blackaby, Mark Robert Albertella et al.*1,352 citations*
33. **[mRNA vaccine-elicited antibodies to SARS-CoV-2 and circulating variants ↗](https://doi.org/10.1038/s41586-021-03324-6)**Zijun Wang, Fabian Schmidt, Yiska Weisblum et al.*1,341 citations*
34. **[Omicron extensively but incompletely escapes Pfizer BNT162b2 neutralization ↗](https://doi.org/10.1038/s41586-021-04387-1)**Sandile Cele, Laurelle Jackson, David S. Khoury et al.*1,331 citations*
35. **[SARS-CoV-2 B.1.617.2 Delta variant replication and immune evasion ↗](https://doi.org/10.1038/s41586-021-03944-y)**Petra Mlčochová, Steven A. Kemp, Mahesh Shanker Dhar et al.*1,330 citations*
36. **[Striking antibody evasion manifested by the Omicron variant of SARS-CoV-2 ↗](https://doi.org/10.1038/s41586-021-04388-0)**Lihong Liu, Sho Iketani, Yicheng Guo et al.*1,286 citations*
37. **[NASH limits anti-tumour surveillance in immunotherapy-treated HCC ↗](https://doi.org/10.1038/s41586-021-03362-0)**Dominik Reinhard Pfister, Nicolás Gonzalo Núñez, Roser Pinyol et al.*1,280 citations*
38. **[Metabolic support of tumour-infiltrating regulatory T cells by lactic acid ↗](https://doi.org/10.1038/s41586-020-03045-2)**McLane J. Watson, Paolo Vignali, Steven J. Mullett et al.*1,259 citations*
39. **[Pattern-recognition receptors are required for NLR-mediated plant immunity ↗](https://doi.org/10.1038/s41586-021-03316-6)**Minhang Yuan, Zeyu Jiang, Guozhi Bi et al.*1,257 citations*
40. **[Self-powered soft robot in the Mariana Trench ↗](https://doi.org/10.1038/s41586-020-03153-z)**Guorui Li, Xiangping Chen, Fanghao Zhou et al.*1,198 citations*
41. **[Shifting attention to accuracy can reduce misinformation online ↗](https://doi.org/10.1038/s41586-021-03344-2)**Gordon Pennycook, Ziv G. Epstein, Mohsen Mosleh et al.*1,139 citations*
42. **[Mutual potentiation of plant immunity by cell-surface and intracellular receptors ↗](https://doi.org/10.1038/s41586-021-03315-7)**Bruno Pok Man Ngou, Hee-Kyung Ahn, Pingtao Ding et al.*1,100 citations*
43. **[SARS-CoV-2 evolution during treatment of chronic infection ↗](https://doi.org/10.1038/s41586-021-03291-y)**Steven A. Kemp, Dami A. Collier, Rawlings P. Datir et al.*1,095 citations*
44. **[Cross-tissue organization of the fibroblast lineage ↗](https://doi.org/10.1038/s41586-021-03549-5)**Matthew B. Buechler, Rachana Pradhan, Akshay T. Krishnamurty et al.*1,087 citations*
45. **[Large-area display textiles integrated with functional systems ↗](https://doi.org/10.1038/s41586-021-03295-8)**Xiang Shi, Yong Zuo, Peng Zhai et al.*1,085 citations*
46. **[Broadly neutralizing antibodies overcome SARS-CoV-2 Omicron antigenic shift ↗](https://doi.org/10.1038/s41586-021-04386-2)**Elisabetta Cameroni, John E. Bowen, Laura E. Rosen et al.*1,080 citations*
47. **[Amazonia as a carbon source linked to deforestation and climate change ↗](https://doi.org/10.1038/s41586-021-03629-6)**Luciana V. Gatti, Luana S. Basso, John Bharat Miller et al.*1,072 citations*
48. **[Bayesian reaction optimization as a tool for chemical synthesis ↗](https://doi.org/10.1038/s41586-021-03213-y)**Benjamin J. Shields, Jason M. Stevens, Jun Li et al.*1,044 citations*
49. **[An integrated space-to-ground quantum communication network over 4,600 kilometres ↗](https://doi.org/10.1038/s41586-020-03093-8)**Yu-Ao Chen, Qiang Zhang, Teng‐Yun Chen et al.*1,040 citations*
50. **[Cell-programmed nutrient partitioning in the tumour microenvironment ↗](https://doi.org/10.1038/s41586-021-03442-1)**Bradley I. Reinfeld, Matthew Z. Madden, Melissa M. Wolf et al.*1,023 citations*
51. **[Skilful precipitation nowcasting using deep generative models of radar ↗](https://doi.org/10.1038/s41586-021-03854-z)**Suman V. Ravuri, Karel Lenc, Matthew Willson et al.*1,006 citations*
52. **[Diverse functional autoantibodies in patients with COVID-19 ↗](https://doi.org/10.1038/s41586-021-03631-y)**Eric Y. Wang, Tianyang Mao, Jon Klein et al.*992 citations*
53. **[Quantum phases of matter on a 256-atom programmable quantum simulator ↗](https://doi.org/10.1038/s41586-021-03582-4)**Sepehr Ebadi, Tout T. Wang, Harry Levine et al.*987 citations*
54. **[Mapping the human genetic architecture of COVID-19 ↗](https://doi.org/10.1038/s41586-021-03767-x)**COVID-19 Host Genetics Initiative, COVID-19 Host Genetics InitiativeLeadership, Mari Niemi et al.*983 citations*
55. **[Exome sequencing and analysis of 454,787 UK Biobank participants ↗](https://doi.org/10.1038/s41586-021-04103-z)**Joshua Backman, Alexander H. Li, Anthony Marcketta et al.*976 citations*
56. **[The power of genetic diversity in genome-wide association studies of lipids ↗](https://doi.org/10.1038/s41586-021-04064-3)**Sarah E. Graham, Shoa L. Clarke, Kuan-Han Wu et al.*967 citations*
57. **[High-performance brain-to-text communication via handwriting ↗](https://doi.org/10.1038/s41586-021-03506-2)**Francis R. Willett, Donald T. Avansino, Leigh R. Hochberg et al.*961 citations*
58. **[An aged immune system drives senescence and ageing of solid organs ↗](https://doi.org/10.1038/s41586-021-03547-7)**Matthew J. Yousefzadeh, Rafael R. Flores, Yi Zhun Zhu et al.*957 citations*
59. **[Unextractable fossil fuels in a 1.5 °C world ↗](https://doi.org/10.1038/s41586-021-03821-8)**Dan Welsby, James Price, Steve Pye et al.*926 citations*
60. **[Increased mortality in community-tested cases of SARS-CoV-2 lineage B.1.1.7 ↗](https://doi.org/10.1038/s41586-021-03426-1)**CMMID COVID-19 Working Group, Nicholas G. Davies, Christopher I Jarvis et al.*922 citations*
61. **[BNT162b2 vaccine induces neutralizing antibodies and poly-specific T cells in humans ↗](https://doi.org/10.1038/s41586-021-03653-6)**Uğur Şahin, Alexander Muik, Isabel Vogler et al.*913 citations*
62. **[The central role of DNA damage in the ageing process ↗](https://doi.org/10.1038/s41586-021-03307-7)**Björn Schumacher, Joris Pothof, Jan Vijg et al.*898 citations*
63. **[Swarm Learning for decentralized and confidential clinical machine learning ↗](https://doi.org/10.1038/s41586-021-03583-3)**Stefanie Warnat‐Herresthal, Hartmut Schultze, Krishnaprasad Shastry et al.*879 citations*
64. **[Structure-based classification of tauopathies ↗](https://doi.org/10.1038/s41586-021-03911-7)**Yang Shi, Wenjuan Zhang, Yang Yang et al.*873 citations*
65. **[Tunable strongly coupled superconductivity in magic-angle twisted trilayer graphene ↗](https://doi.org/10.1038/s41586-021-03192-0)**Jeong Min Park, Yuan Cao, Kenji Watanabe et al.*855 citations*
66. **[High-resolution X-ray luminescence extension imaging ↗](https://doi.org/10.1038/s41586-021-03251-6)**Xiangyu Ou, Xian Qin, Bolong Huang et al.*853 citations*
67. **[NINJ1 mediates plasma membrane rupture during lytic cell death ↗](https://doi.org/10.1038/s41586-021-03218-7)**Nobuhiko Kayagaki, Opher S. Kornfeld, Bettina L. Lee et al.*838 citations*
68. **[Leading hadronic contribution to the muon magnetic moment from lattice QCD ↗](https://doi.org/10.1038/s41586-021-03418-1)**Sz. Borsányi, Z. Fodor, Jana N. Guenther et al.*824 citations*
69. **[Ligand-engineered bandgap stability in mixed-halide perovskite LEDs ↗](https://doi.org/10.1038/s41586-021-03217-8)**Yasser Hassan, Jong Hyun Park, Michael L. Crawford et al.*824 citations*
70. **[SARS-CoV-2 mRNA vaccines induce persistent human germinal centre responses ↗](https://doi.org/10.1038/s41586-021-03738-2)**Jackson S. Turner, Jane A. O’Halloran, Elizaveta Kalaidina et al.*824 citations*
71. **[A neuroanatomical basis for electroacupuncture to drive the vagal–adrenal axis ↗](https://doi.org/10.1038/s41586-021-04001-4)**Shenbin Liu, Zhifu Wang, Yang‐Shuai Su et al.*823 citations*
72. **[Direct observation of chemical short-range order in a medium-entropy alloy ↗](https://doi.org/10.1038/s41586-021-03428-z)**Xuefei Chen, Qi S. Wang, Zhiying Cheng et al.*819 citations*
73. **[Protecting the global ocean for biodiversity, food and climate ↗](https://doi.org/10.1038/s41586-021-03371-z)**Enric Sala, Juan Mayorga, Darcy Bradley et al.*811 citations*
74. **[Half a century of global decline in oceanic sharks and rays ↗](https://doi.org/10.1038/s41586-020-03173-9)**Nathan Pacoureau, Cassandra L. Rigby, Peter M. Kyne et al.*808 citations*
75. **[Comparative cellular analysis of motor cortex in human, marmoset and mouse ↗](https://doi.org/10.1038/s41586-021-03465-8)**Trygve E. Bakken, Nikolas L. Jorstad, Qiwen Hu et al.*805 citations*
76. **[COVID-19 tissue atlases reveal SARS-CoV-2 pathology and cellular targets ↗](https://doi.org/10.1038/s41586-021-03570-8)**Toni Marie Delorey, Carly G.K. Ziegler, Graham Heimberg et al.*794 citations*
77. **[Distinct fission signatures predict mitochondrial degradation or biogenesis ↗](https://doi.org/10.1038/s41586-021-03510-6)**Tatjana Kleele, Timo Rey, Julius Winter et al.*787 citations*
78. **[In vivo CRISPR base editing of PCSK9 durably lowers cholesterol in primates ↗](https://doi.org/10.1038/s41586-021-03534-y)**Kiran Musunuru, Alexandra C. Chadwick, Taiji Mizoguchi et al.*785 citations*
79. **[AIM2 forms a complex with pyrin and ZBP1 to drive PANoptosis and host defence ↗](https://doi.org/10.1038/s41586-021-03875-8)**SangJoon Lee, Rajendra Karki, Yaqiu Wang et al.*783 citations*
80. **[Disease variant prediction with deep generative models of evolutionary data ↗](https://doi.org/10.1038/s41586-021-04043-8)**Jonathan Frazer, Pascal Notin, Mafalda Dias et al.*782 citations*
81. **[Closed-loop recycling of polyethylene-like materials ↗](https://doi.org/10.1038/s41586-020-03149-9)**Manuel Häußler, Marcel Eck, Dario Rothauer et al.*752 citations*
82. **[Distribution control enables efficient reduced-dimensional perovskite LEDs ↗](https://doi.org/10.1038/s41586-021-03997-z)**Dongxin Ma, Kebin Lin, Yitong Dong et al.*752 citations*
83. **[A lymphocyte–microglia–astrocyte axis in chronic active multiple sclerosis ↗](https://doi.org/10.1038/s41586-021-03892-7)**Martina Absinta, Dragan Maric, Marjan Gharagozloo et al.*751 citations*
84. **[Age-related immune response heterogeneity to SARS-CoV-2 vaccine BNT162b2 ↗](https://doi.org/10.1038/s41586-021-03739-1)**Dami A. Collier, Isabella A.T.M. Ferreira, Prasanti Kotagiri et al.*747 citations*
85. **[A dynamic stability design strategy for lithium metal solid state batteries ↗](https://doi.org/10.1038/s41586-021-03486-3)**Luhan Ye, Xin Li*739 citations*
86. **[MicroRNA sequence codes for small extracellular vesicle release and cellular retention ↗](https://doi.org/10.1038/s41586-021-04234-3)**Rubén García-Martín, Guoxiao Wang, Bruna Brasil Brandão et al.*731 citations*
87. **[Circuits between infected macrophages and T cells in SARS-CoV-2 pneumonia ↗](https://doi.org/10.1038/s41586-020-03148-w)**Rogan A. Grant, Luisa Morales‐Nebreda, Nikolay S. Markov et al.*727 citations*
88. **[Naturally enhanced neutralizing breadth against SARS-CoV-2 one year after infection ↗](https://doi.org/10.1038/s41586-021-03696-9)**Zijun Wang, Frauke Muecksch, Dennis J. Schaefer-Babajew et al.*721 citations*
89. **[The KEYNOTE-811 trial of dual PD-1 and HER2 blockade in HER2-positive gastric cancer ↗](https://doi.org/10.1038/s41586-021-04161-3)**Yelena Yuriy Janjigian, Akihito Kawazoe, Patricio Eduardo Yanez et al.*720 citations*
90. **[Cells of the human intestinal tract mapped across space and time ↗](https://doi.org/10.1038/s41586-021-03852-1)**Rasa Elmentaite, Natsuhiko Kumasaka, Kenny Roberts et al.*699 citations*
91. **[Correlative operando microscopy of oxygen evolution electrocatalysts ↗](https://doi.org/10.1038/s41586-021-03454-x)**J. Tyler Mefford, Andrew R. Akbashev, Minkyung Kang et al.*699 citations*
92. **[Microplastics are everywhere — but are they harmful? ↗](https://doi.org/10.1038/d41586-021-01143-3)**XiaoZhi Lim*697 citations*
93. **[Loss of furin cleavage site attenuates SARS-CoV-2 pathogenesis ↗](https://doi.org/10.1038/s41586-021-03237-4)**Bryan A. Johnson, Xuping Xie, Adam L. Bailey et al.*693 citations*
94. **[BNT162b vaccines protect rhesus macaques from SARS-CoV-2 ↗](https://doi.org/10.1038/s41586-021-03275-y)**Annette B. Vogel, Isis Kanevsky, Ye Che et al.*691 citations*
95. **[Thermal-expansion offset for high-performance fuel cell cathodes ↗](https://doi.org/10.1038/s41586-021-03264-1)**Yuan Zhang, Bin Chen, Daqin Guan et al.*691 citations*
96. **[Widespread deoxygenation of temperate lakes ↗](https://doi.org/10.1038/s41586-021-03550-y)**Stephen F. Jane, Gretchen J. A. Hansen, Benjamin Michael Kraemer et al.*690 citations*
97. **[A trade-off between plant and soil carbon storage under elevated CO2 ↗](https://doi.org/10.1038/s41586-021-03306-8)**César Terrer, Richard P. Phillips, Bruce A. Hungate et al.*687 citations*
98. **[Multilevel proteomics reveals host perturbations by SARS-CoV-2 and SARS-CoV ↗](https://doi.org/10.1038/s41586-021-03493-4)**Alexey Stukalov, Virginie Girault, Vincent Grass et al.*685 citations*
99. **[Quantum simulation of 2D antiferromagnets with hundreds of Rydberg atoms ↗](https://doi.org/10.1038/s41586-021-03585-1)**Pascal Scholl, Michael Schuler, Hannah J. Williams et al.*685 citations*
100. **[A molecular single-cell lung atlas of lethal COVID-19 ↗](https://doi.org/10.1038/s41586-021-03569-1)**Johannes C. Melms, Jana Biermann, Huachao Huang et al.*679 citations*

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