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title: "Nature 2021 — all accepted papers, page 3 — the most cited first"
description: "All 3,641 papers of Nature 2021 · page 3 of 37, the most cited first; those with our summary link to it."
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# Nature 2021: all accepted papers

3,641 papers · page 3 of 37 · the most cited first · [← Nature 2021](https://md.research.elamur.ai/free/en/venues/nature/2021/)

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201. **[Lipid signalling enforces functional specialization of Treg cells in tumours ↗](https://doi.org/10.1038/s41586-021-03235-6)**Seon Ah Lim, Jun Wei, Thanh-Long M. Nguyen et al.*426 citations*
202. **[The universal visitation law of human mobility ↗](https://doi.org/10.1038/s41586-021-03480-9)**Markus Schläpfer, Lei Dong, Kevin P. O’Keeffe et al.*425 citations*
203. **[Meningeal lymphatics affect microglia responses and anti-Aβ immunotherapy ↗](https://doi.org/10.1038/s41586-021-03489-0)**Sandro Dá Mesquita, Zachary Papadopoulos, Taitea Dykstra et al.*423 citations*
204. **[The role of retrotransposable elements in ageing and age-associated diseases ↗](https://doi.org/10.1038/s41586-021-03542-y)**Vera A. Gorbunova, Andrei Seluanov, Paolo Mita et al.*423 citations*
205. **[Spread of a SARS-CoV-2 variant through Europe in the summer of 2020 ↗](https://doi.org/10.1038/s41586-021-03677-y)**Emma B. Hodcroft, Moira Zuber, Sarah Nadeau et al.*422 citations*
206. **[Exercise plasma boosts memory and dampens brain inflammation via clusterin ↗](https://doi.org/10.1038/s41586-021-04183-x)**Zurine D. Miguel, Nathalie Khoury, Michael J. Betley et al.*418 citations*
207. **[Integrated spatial genomics reveals global architecture of single nuclei ↗](https://doi.org/10.1038/s41586-020-03126-2)**Yodai Takei, Jina Yun, Shiwei Zheng et al.*418 citations*
208. **[Morphological diversity of single neurons in molecularly defined cell types ↗](https://doi.org/10.1038/s41586-021-03941-1)**Hanchuan Peng, Peng Xie, Lijuan Liu et al.*416 citations*
209. **[Regulatory genomic circuitry of human disease loci by integrative epigenomics ↗](https://doi.org/10.1038/s41586-020-03145-z)**Carles A. Boix, Benjamin T. James, Yongjin Park et al.*416 citations*
210. **[Origins of modern human ancestry ↗](https://doi.org/10.1038/s41586-021-03244-5)**Anders Bergström, Chris B. Stringer, Mateja Hajdinjak et al.*415 citations*
211. **[A quantum enhanced search for dark matter axions ↗](https://doi.org/10.1038/s41586-021-03226-7)**K. M. Backes, D. A. Palken, S. Al Kenany et al.*414 citations*
212. **[Lessons from the host defences of bats, a unique viral reservoir ↗](https://doi.org/10.1038/s41586-020-03128-0)**Aaron Trent Irving, Matae Ahn, Geraldine Goh et al.*413 citations*
213. **[The emergence, genomic diversity and global spread of SARS-CoV-2 ↗](https://doi.org/10.1038/s41586-021-04188-6)**Juan Li, Shengjie Lai, George F. Gao et al.*413 citations*
214. **[A metabolomics pipeline for the mechanistic interrogation of the gut microbiome ↗](https://doi.org/10.1038/s41586-021-03707-9)**Shuo Han, Will Van Treuren, Curt R. Fischer et al.*409 citations*
215. **[SARS-CoV-2 infection is effectively treated and prevented by EIDD-2801 ↗](https://doi.org/10.1038/s41586-021-03312-w)**Angela R. Wahl, Lisa E. Gralinski, Claire E. Johnson et al.*409 citations*
216. **[Structured fabrics with tunable mechanical properties ↗](https://doi.org/10.1038/s41586-021-03698-7)**Yifan Wang, Liuchi Li, Douglas C. Hofmann et al.*407 citations*
217. **[Connectomes across development reveal principles of brain maturation ↗](https://doi.org/10.1038/s41586-021-03778-8)**Daniel K. Witvliet, Ben Mulcahy, James Kameron Mitchell et al.*406 citations*
218. **[A four-qubit germanium quantum processor ↗](https://doi.org/10.1038/s41586-021-03332-6)**Nico W. Hendrickx, William I. L. Lawrie, Maximilian Russ et al.*405 citations*
219. **[Pre-existing polymerase-specific T cells expand in abortive seronegative SARS-CoV-2 ↗](https://doi.org/10.1038/s41586-021-04186-8)**Leo Swadling, Mariana O. Diniz, Nathalie M. Schmidt et al.*405 citations*
220. **[Synthon-based ligand discovery in virtual libraries of over 11 billion compounds ↗](https://doi.org/10.1038/s41586-021-04220-9)**Arman A. Sadybekov, Anastasiia Sadybekov, Yongfeng Liu et al.*405 citations*
221. **[The origins and spread of domestic horses from the Western Eurasian steppes ↗](https://doi.org/10.1038/s41586-021-04018-9)**Pablo Librado, Naveed Khan, Antoine Fages et al.*405 citations*
222. **[Drop-in fuels from sunlight and air ↗](https://doi.org/10.1038/s41586-021-04174-y)**Remo Schäppi, David Rutz, Fabian Dähler et al.*404 citations*
223. **[Drugs that inhibit TMEM16 proteins block SARS-CoV-2 spike-induced syncytia ↗](https://doi.org/10.1038/s41586-021-03491-6)**Luca Braga, Hashim Ali, Ilaria Secco et al.*403 citations*
224. **[SARS-CoV-2 infection in free-ranging white-tailed deer ↗](https://doi.org/10.1038/s41586-021-04353-x)**Vanessa L. Hale, Patricia M. Dennis, Dillon S. McBride et al.*403 citations*
225. **[Single-crystal, large-area, fold-free monolayer graphene ↗](https://doi.org/10.1038/s41586-021-03753-3)**Meihui Wang, Ming Yang Huang, Da Luo et al.*401 citations*
226. **[Tumour DDR1 promotes collagen fibre alignment to instigate immune exclusion ↗](https://doi.org/10.1038/s41586-021-04057-2)**Xiujie Sun, Bogang Wu, Huai-Chin Chiang et al.*401 citations*
227. **[Skeletal editing through direct nitrogen deletion of secondary amines ↗](https://doi.org/10.1038/s41586-021-03448-9)**Sean H. Kennedy, Balu D. Dherange, Kathleen J. Berger et al.*399 citations*
228. **[A universal 3D imaging sensor on a silicon photonics platform ↗](https://doi.org/10.1038/s41586-021-03259-y)**Christopher M. Rogers, Alexander Y. Piggott, David J. Thomson et al.*398 citations*
229. **[Ubiquitylation of lipopolysaccharide by RNF213 during bacterial infection ↗](https://doi.org/10.1038/s41586-021-03566-4)**Elsje G. Otten, Emma Werner, Ana Crespillo-Casado et al.*394 citations*
230. **[CTLA-4 blockade drives loss of Treg stability in glycolysis-low tumours ↗](https://doi.org/10.1038/s41586-021-03326-4)**Roberta Zappasodi, Inna S. Serganova, Ivan Cohen et al.*393 citations*
231. **[Global potential for harvesting drinking water from air using solar energy ↗](https://doi.org/10.1038/s41586-021-03900-w)**Jackson Lord, Ashley Thomas, Neil D. Treat et al.*393 citations*
232. **[Shared mechanisms underlie the control of working memory and attention ↗](https://doi.org/10.1038/s41586-021-03390-w)**Matthew F. Panichello, Timothy J. Buschman*393 citations*
233. **[A transcriptomic and epigenomic cell atlas of the mouse primary motor cortex ↗](https://doi.org/10.1038/s41586-021-03500-8)**Zizhen Yao, Hanqing Liu, Fangming Xie et al.*391 citations*
234. **[Unravelling the collateral damage of antibiotics on gut bacteria ↗](https://doi.org/10.1038/s41586-021-03986-2)**Lisa Maier, Camille Goemans, Jakob Wirbel et al.*391 citations*
235. **[Frequency ratio measurements at 18-digit accuracy using an optical clock network ↗](https://doi.org/10.1038/s41586-021-03253-4)**K. Beloy, Martha I. Bodine, Tobias Bothwell et al.*389 citations*
236. **[Rapid and stable mobilization of CD8+ T cells by SARS-CoV-2 mRNA vaccine ↗](https://doi.org/10.1038/s41586-021-03841-4)**Valerie Oberhardt, Hendrik Luxenburger, Janine Sophie Kemming et al.*389 citations*
237. **[Base editing of haematopoietic stem cells rescues sickle cell disease in mice ↗](https://doi.org/10.1038/s41586-021-03609-w)**Gregory A. Newby, Jonathan S. Yen, Kaitly J. Woodard et al.*386 citations*
238. **[Aged skeletal stem cells generate an inflammatory degenerative niche ↗](https://doi.org/10.1038/s41586-021-03795-7)**Thomas H. Ambrosi, Owen Marecic, Adrian McArdle et al.*384 citations*
239. **[Fine-root traits in the global spectrum of plant form and function ↗](https://doi.org/10.1038/s41586-021-03871-y)**Carlos P. Carmona, Carlos Guillermo Bueno, Aurèle Toussaint et al.*384 citations*
240. **[Exponential suppression of bit or phase errors with cyclic error correction ↗](https://doi.org/10.1038/s41586-021-03588-y)**Google Quantum AI, Zijun Chen, Kevin J. Satzinger et al.*381 citations*
241. **[Real-time optimal quantum control of mechanical motion at room temperature ↗](https://doi.org/10.1038/s41586-021-03602-3)**Lorenzo Magrini, Philipp Rosenzweig, Constanze Bach et al.*381 citations*
242. **[Estimating a social cost of carbon for global energy consumption ↗](https://doi.org/10.1038/s41586-021-03883-8)**Ashwin Rode, Tamma A. Carleton, Michael Delgado et al.*380 citations*
243. **[Imaging two-dimensional generalized Wigner crystals ↗](https://doi.org/10.1038/s41586-021-03874-9)**Hongyuan Li, Shaowei Li, Emma C. Regan et al.*379 citations*
244. **[The mutational landscape of human somatic and germline cells ↗](https://doi.org/10.1038/s41586-021-03822-7)**Luiza Moore, Alex Cagan, Tim H. H. Coorens et al.*377 citations*
245. **[TIM-3 restrains anti-tumour immunity by regulating inflammasome activation ↗](https://doi.org/10.1038/s41586-021-03626-9)**Karen Olivia Dixon, Marcin Tabaka, Markus A. Schramm et al.*372 citations*
246. **[METTL3 regulates heterochromatin in mouse embryonic stem cells ↗](https://doi.org/10.1038/s41586-021-03210-1)**Wenqi Xu, Jiahui Li, Chenxi He et al.*371 citations*
247. **[Direct radiative effects of airborne microplastics ↗](https://doi.org/10.1038/s41586-021-03864-x)**Laura E. Revell, Peter Kuma, Eric C. Le Ru et al.*370 citations*
248. **[Evolution of enhanced innate immune evasion by SARS-CoV-2 ↗](https://doi.org/10.1038/s41586-021-04352-y)**Lucy Thorne, Mehdi Bouhaddou, Ann‐Kathrin Reuschl et al.*370 citations*
249. **[Direct imaging of single-molecule electrochemical reactions in solution ↗](https://doi.org/10.1038/s41586-021-03715-9)**Jinrun Dong, Yuxian Lu, Yang Xu et al.*369 citations*
250. **[In situ formation of ZnOx species for efficient propane dehydrogenation ↗](https://doi.org/10.1038/s41586-021-03923-3)**Dan Zhao, Xinxin Tian, Dmitry E. Doronkin et al.*368 citations*
251. **[Polypeptide organic radical batteries ↗](https://doi.org/10.1038/s41586-021-03399-1)**Tan P. Nguyen, Alexandra Danielle Easley, Na‐Ri Kang et al.*368 citations*
252. **[Spatiotemporal origin of soil water taken up by vegetation ↗](https://doi.org/10.1038/s41586-021-03958-6)**Gonzalo Miguez‐Macho, Ying Fan*367 citations*
253. **[Epigenetic silencing by SETDB1 suppresses tumour intrinsic immunogenicity ↗](https://doi.org/10.1038/s41586-021-03520-4)**Gabriel K. Griffin, Jingyi Wu, Arvin Iracheta‐Vellve et al.*365 citations*
254. **[Local and systemic responses to SARS-CoV-2 infection in children and adults ↗](https://doi.org/10.1038/s41586-021-04345-x)**Masahiro Yoshida, Kaylee B. Worlock, Ni Huang et al.*364 citations*
255. **[The spatial landscape of lung pathology during COVID-19 progression ↗](https://doi.org/10.1038/s41586-021-03475-6)**André F. Rendeiro, Hiranmayi Ravichandran, Yaron Bram et al.*364 citations*
256. **[Evidence for unconventional superconductivity in twisted bilayer graphene ↗](https://doi.org/10.1038/s41586-021-04121-x)**Myungchul Oh, Kevin P. Nuckolls, Dillon Wong et al.*362 citations*
257. **[Integrating explanation and prediction in computational social science ↗](https://doi.org/10.1038/s41586-021-03659-0)**Jake M. Hofman, Duncan J. Watts, Susan Carleton Athey et al.*361 citations*
258. **[Modelling human blastocysts by reprogramming fibroblasts into iBlastoids ↗](https://doi.org/10.1038/s41586-021-03372-y)**Xiaodong Liu, Jia Ping Tan, Jan Schröder et al.*361 citations*
259. **[Transposon-associated TnpB is a programmable RNA-guided DNA endonuclease ↗](https://doi.org/10.1038/s41586-021-04058-1)**Tautvydas Karvelis, Gytis Druteika, Greta Bigelyte et al.*359 citations*
260. **[Activation of TIR signalling boosts pattern-triggered immunity ↗](https://doi.org/10.1038/s41586-021-03987-1)**Hainan Tian, Zhongshou Wu, Siyu Chen et al.*358 citations*
261. **[Multi-kingdom ecological drivers of microbiota assembly in preterm infants ↗](https://doi.org/10.1038/s41586-021-03241-8)**Chitong Rao, Katharine Z. Coyte, Wayne Bainter et al.*356 citations*
262. **[The RNA m6A reader YTHDC1 silences retrotransposons and guards ES cell identity ↗](https://doi.org/10.1038/s41586-021-03313-9)**Jiadong Liu, Mingwei Gao, Jiangping He et al.*356 citations*
263. **[Seasonal origin of the thermal maxima at the Holocene and the last interglacial ↗](https://doi.org/10.1038/s41586-020-03155-x)**Samantha C. Bova, Yair Rosenthal, Zhengyu Liu et al.*355 citations*
264. **[ecDNA hubs drive cooperative intermolecular oncogene expression ↗](https://doi.org/10.1038/s41586-021-04116-8)**King L. Hung, Kathryn E. Yost, Liangqi Xie et al.*354 citations*
265. **[Bifunctional nanoprecipitates strengthen and ductilize a medium-entropy alloy ↗](https://doi.org/10.1038/s41586-021-03607-y)**Ying Yang, Tianyi Chen, Lizhen Tan et al.*351 citations*
266. **[Microbiota regulate social behaviour via stress response neurons in the brain ↗](https://doi.org/10.1038/s41586-021-03669-y)**Wei‐Li Wu, Mark D. Adame, Chia-Wei Liou et al.*349 citations*
267. **[Virus-induced senescence is a driver and therapeutic target in COVID-19 ↗](https://doi.org/10.1038/s41586-021-03995-1)**Soyoung Lee, Yong Yu, Jakob Trimpert et al.*348 citations*
268. **[A transcriptomic atlas of mouse cerebellar cortex comprehensively defines cell types ↗](https://doi.org/10.1038/s41586-021-03220-z)**Velina Kozareva, Caroline Martin, Tomás Osorno et al.*347 citations*
269. **[Free-electron lasing at 27 nanometres based on a laser wakefield accelerator ↗](https://doi.org/10.1038/s41586-021-03678-x)**Wentao Wang, Ke Feng, Lintong Ke et al.*347 citations*
270. **[Layer Hall effect in a 2D topological axion antiferromagnet ↗](https://doi.org/10.1038/s41586-021-03679-w)**Anyuan Gao, Yu-Fei Liu, Chaowei Hu et al.*347 citations*
271. **[Five reasons why COVID herd immunity is probably impossible ↗](https://doi.org/10.1038/d41586-021-00728-2)**Christie Aschwanden*346 citations*
272. **[Nature-based solutions can help cool the planet — if we act now ↗](https://doi.org/10.1038/d41586-021-01241-2)**Cecile A. J. Girardin, Stuart Jenkins, Nathalie Seddon A et al.*345 citations*
273. **[Quantum control of a nanoparticle optically levitated in cryogenic free space ↗](https://doi.org/10.1038/s41586-021-03617-w)**Felix Tebbenjohanns, M. Luisa Mattana, Massimiliano Rossi et al.*345 citations*
274. **[Quantum-enhanced nonlinear microscopy ↗](https://doi.org/10.1038/s41586-021-03528-w)**Catxere A. Casacio, Lars S. Madsen, Alex Terrasson et al.*344 citations*
275. **[Fault-tolerant control of an error-corrected qubit ↗](https://doi.org/10.1038/s41586-021-03928-y)**Laird Egan, Dripto M. Debroy, Crystal Noel et al.*343 citations*
276. **[Antiviral activity of bacterial TIR domains via immune signalling molecules ↗](https://doi.org/10.1038/s41586-021-04098-7)**Gal Ofir, Ehud Herbst, Maya Baroz et al.*342 citations*
277. **[Million-year-old DNA sheds light on the genomic history of mammoths ↗](https://doi.org/10.1038/s41586-021-03224-9)**Tom van der Valk, Patrícia Pečnerová, David Díez‐del‐Molino et al.*342 citations*
278. **[BNT162b2-elicited neutralization of B.1.617 and other SARS-CoV-2 variants ↗](https://doi.org/10.1038/s41586-021-03693-y)**Jianying Liu, Yang Liu, Hongjie Xia et al.*341 citations*
279. **[The structure, function and evolution of a complete human chromosome 8 ↗](https://doi.org/10.1038/s41586-021-03420-7)**Glennis A. Logsdon, Mitchell Robert Vollger, PingHsun Hsieh et al.*341 citations*
280. **[Loop extrusion as a mechanism for formation of DNA damage repair foci ↗](https://doi.org/10.1038/s41586-021-03193-z)**Coline Arnould, Vincent Rocher, Anne-Laure Finoux et al.*339 citations*
281. **[Hepatic stellate cells suppress NK cell-sustained breast cancer dormancy ↗](https://doi.org/10.1038/s41586-021-03614-z)**Ana Luísa Correia, Joao C. Guimaraes, Priska Auf der Maur et al.*337 citations*
282. **[Extremely anisotropic van der Waals thermal conductors ↗](https://doi.org/10.1038/s41586-021-03867-8)**Shi En Kim, Fauzia Mujid, Akash Rai et al.*335 citations*
283. **[Half- and quarter-metals in rhombohedral trilayer graphene ↗](https://doi.org/10.1038/s41586-021-03938-w)**Haoxin Zhou, Tian Xie, Areg Ghazaryan et al.*334 citations*
284. **[Origins of structural and electronic transitions in disordered silicon ↗](https://doi.org/10.1038/s41586-020-03072-z)**Volker L. Deringer, Noam Bernstein, Gábor Cśanyi et al.*334 citations*
285. **[Time-crystalline eigenstate order on a quantum processor ↗](https://doi.org/10.1038/s41586-021-04257-w)**Xiao Mi, Matteo Ippoliti, Chris Quintana et al.*334 citations*
286. **[Functional HPV-specific PD-1+ stem-like CD8 T cells in head and neck cancer ↗](https://doi.org/10.1038/s41586-021-03862-z)**Christiane Sigrid Eberhardt, Haydn Thomas Kissick, Mihir R. Patel et al.*333 citations*
287. **[Summertime increases in upper-ocean stratification and mixed-layer depth ↗](https://doi.org/10.1038/s41586-021-03303-x)**Jean‐Baptiste Sallée, Violaine Pellichero, Camille Hayatte Akhoudas et al.*333 citations*
288. **[Transition metal-catalysed molecular n-doping of organic semiconductors ↗](https://doi.org/10.1038/s41586-021-03942-0)**Han Guo, Chi‐Yuan Yang, Xianhe Zhang et al.*333 citations*
289. **[High-entropy polymer produces a giant electrocaloric effect at low fields ↗](https://doi.org/10.1038/s41586-021-04189-5)**Xiaoshi Qian, Donglin Han, Lirong Zheng et al.*332 citations*
290. **[Geometry of abstract learned knowledge in the hippocampus ↗](https://doi.org/10.1038/s41586-021-03652-7)**Edward H. Nieh, Manuel Schottdorf, Nicolas W. Freeman et al.*331 citations*
291. **[Continuous Mott transition in semiconductor moire superlattices ↗](https://doi.org/10.1038/s41586-021-03853-0)**Tingxin Li, Shengwei Jiang, Lizhong Li et al.*330 citations*
292. **[Gut-licensed IFNγ+ NK cells drive LAMP1+TRAIL+ anti-inflammatory astrocytes ↗](https://doi.org/10.1038/s41586-020-03116-4)**Liliana María Sanmarco, Michael A. Wheeler, Cristina Gutiérrez‐Vázquez et al.*330 citations*
293. **[Bioaccumulation of therapeutic drugs by human gut bacteria ↗](https://doi.org/10.1038/s41586-021-03891-8)**Martina Klünemann, Sergej Andrejev, Sonja Blasche et al.*327 citations*
294. **[Adjuvanting a subunit COVID-19 vaccine to induce protective immunity ↗](https://doi.org/10.1038/s41586-021-03530-2)**Prabhu S. Arunachalam, Alexandra C. Walls, Nadia Golden et al.*326 citations*
295. **[Agrochemicals interact synergistically to increase bee mortality ↗](https://doi.org/10.1038/s41586-021-03787-7)**Harry Siviter, Emily J. Bailes, Callum D. Martin et al.*326 citations*
296. **[Biogeochemical extremes and compound events in the ocean ↗](https://doi.org/10.1038/s41586-021-03981-7)**Nicolas Gruber, Philip W. Boyd, Thomas Lukas Frölicher et al.*326 citations*
297. **[Towards a rigorous understanding of societal responses to climate change ↗](https://doi.org/10.1038/s41586-021-03190-2)**Dagomar Degroot, Kevin John Anchukaitis, Martin Bauch et al.*326 citations*
298. **[Nuclear sensing of breaks in mitochondrial DNA enhances immune surveillance ↗](https://doi.org/10.1038/s41586-021-03269-w)**Marco Tigano, Danielle Vargas, Samuel Tremblay‐Belzile et al.*325 citations*
299. **[Expanded diversity of Asgard archaea and their relationships with eukaryotes ↗](https://doi.org/10.1038/s41586-021-03494-3)**Yang Liu, Kira S. Makarova, Wen-Cong Huang et al.*324 citations*
300. **[Measuring human capital using global learning data ↗](https://doi.org/10.1038/s41586-021-03323-7)**Noam Angrist, Simeon Djankov, Pinelopi Koujianou Goldberg et al.*323 citations*

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