---
title: "Nature 2021 — all accepted papers, page 6 — the most cited first"
description: "All 3,641 papers of Nature 2021 · page 6 of 37, the most cited first; those with our summary link to it."
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# Nature 2021: all accepted papers

3,641 papers · page 6 of 37 · the most cited first · [← Nature 2021](https://md.research.elamur.ai/free/en/venues/nature/2021/)

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501. **[Considerable escape of SARS-CoV-2 Omicron to antibody neutralization ↗](https://doi.org/10.1038/d41586-021-03827-2)**Delphine Planas, Nell Saunders, Piet Maes et al.*197 citations*
502. **[Localization atomic force microscopy ↗](https://doi.org/10.1038/s41586-021-03551-x)**George R. Heath, Ekaterina D. Kots, Janice Robertson et al.*197 citations*
503. **[A pan-serotype dengue virus inhibitor targeting the NS3–NS4B interaction ↗](https://doi.org/10.1038/s41586-021-03990-6)**Suzanne J. F. Kaptein, Olivia Goethals, Dominik Kiemel et al.*196 citations*
504. **[Single-defect phonons imaged by electron microscopy ↗](https://doi.org/10.1038/s41586-020-03049-y)**Xingxu Yan, Chengyan Liu, Chaitanya Avinash Gadre et al.*196 citations*
505. **[The genomic origins of the Bronze Age Tarim Basin mummies ↗](https://doi.org/10.1038/s41586-021-04052-7)**Fan Zhang, Chao Ning, Ashley Scott et al.*195 citations*
506. **[Uncovering global-scale risks from commercial chemicals in air ↗](https://doi.org/10.1038/s41586-021-04134-6)**Qifan Liu, Li Li, Xianming Zhang et al.*195 citations*
507. **[Three-dimensional nanoprinting via charged aerosol jets ↗](https://doi.org/10.1038/s41586-021-03353-1)**Wooik Jung, Yoon-ho Jung, Peter V. Pikhitsa et al.*194 citations*
508. **[Untangling introductions and persistence in COVID-19 resurgence in Europe ↗](https://doi.org/10.1038/s41586-021-03754-2)**Philippe Lemey, Nick Ruktanonchai, Samuel Leandro Hong et al.*194 citations*
509. **[Aziridine synthesis by coupling amines and alkenes via an electrogenerated dication ↗](https://doi.org/10.1038/s41586-021-03717-7)**Dylan E. Holst, Diana J. Wang, Min Ji Kim et al.*193 citations*
510. **[Shigella evades pyroptosis by arginine ADP-riboxanation of caspase-11 ↗](https://doi.org/10.1038/s41586-021-04020-1)**Zilin Li, Wang Liu, Jiaqi Fu et al.*193 citations*
511. **[Mutant clones in normal epithelium outcompete and eliminate emerging tumours ↗](https://doi.org/10.1038/s41586-021-03965-7)**Bartomeu Colom, Albert Herms, Michael Hall et al.*192 citations*
512. **[Patterns of de novo tandem repeat mutations and their role in autism ↗](https://doi.org/10.1038/s41586-020-03078-7)**Ileena Mitra, Bonnie B. Huang, Nima Mousavi et al.*192 citations*
513. **[A radical approach for the selective C–H borylation of azines ↗](https://doi.org/10.1038/s41586-021-03637-6)**Ji Hye Kim, Timothée Constantin, Marco Simonetti et al.*191 citations*
514. **[How deregulation, drought and increasing fire impact Amazonian biodiversity ↗](https://doi.org/10.1038/s41586-021-03876-7)**Xiao Feng, Cory Merow, Zhihua Liu et al.*191 citations*
515. **[RETRACTED ARTICLE: A 10 per cent increase in global land evapotranspiration from 2003 to 2019 ↗](https://doi.org/10.1038/s41586-021-03503-5)**Madeleine Pascolini‐Campbell, John Thomas Reager, Hrishikesh Arvind Chandanpurkar et al.*191 citations*
516. **[UTX condensation underlies its tumour-suppressive activity ↗](https://doi.org/10.1038/s41586-021-03903-7)**Bi Shi, Wei Li, Yansu Song et al.*191 citations*
517. **[Beyond Omicron: what’s next for COVID’s viral evolution ↗](https://doi.org/10.1038/d41586-021-03619-8)**Ewen Callaway*190 citations*
518. **[An open-access volume electron microscopy atlas of whole cells and tissues ↗](https://doi.org/10.1038/s41586-021-03992-4)**C. Shan Xu, Song Pang, Gleb E. Shtengel et al.*189 citations*
519. **[Burden and characteristics of COVID-19 in the United States during 2020 ↗](https://doi.org/10.1038/s41586-021-03914-4)**Sen Pei, Teresa K. Yamana, Sasikiran Kandula et al.*189 citations*
520. **[Convergent somatic mutations in metabolism genes in chronic liver disease ↗](https://doi.org/10.1038/s41586-021-03974-6)**Stanley Ng, Foad J. Rouhani, Simon F. Brunner et al.*189 citations*
521. **[High CO2 levels drive the TCA cycle backwards towards autotrophy ↗](https://doi.org/10.1038/s41586-021-03456-9)**Lydia Steffens, Eugenio Pettinato, Thomas M. Steiner et al.*189 citations*
522. **[Pandemic burnout is rampant in academia ↗](https://doi.org/10.1038/d41586-021-00663-2)**Virginia L. Gewin*189 citations*
523. **[A solar C/O and sub-solar metallicity in a hot Jupiter atmosphere ↗](https://doi.org/10.1038/s41586-021-03912-6)**Michael R. Line, Matteo Brogi, Jacob L. Bean et al.*188 citations*
524. **[Thinking clearly about social aspects of infectious disease transmission ↗](https://doi.org/10.1038/s41586-021-03694-x)**Caroline O. Buckee, Abdisalan Mohamed Noor, Lisa Sattenspiel*188 citations*
525. **[cGAS-like receptors sense RNA and control 3′2′-cGAMP signalling in Drosophila ↗](https://doi.org/10.1038/s41586-021-03743-5)**Kailey M. Slavik, Benjamin R. Morehouse, Adelyn E. Ragucci et al.*188 citations*
526. **[In-orbit demonstration of an iodine electric propulsion system ↗](https://doi.org/10.1038/s41586-021-04015-y)**Dmytro Rafalskyi, Javier Martínez Martínez, Lui Txai Calvoso Habl et al.*187 citations*
527. **[Rapid microbial methanogenesis during CO2 storage in hydrocarbon reservoirs ↗](https://doi.org/10.1038/s41586-021-04153-3)**Rebecca Tyne, Peter H. Barry, Michael J. Lawson et al.*187 citations*
528. **[Ubiquitin ligation to F-box protein targets by SCF–RBR E3–E3 super-assembly ↗](https://doi.org/10.1038/s41586-021-03197-9)**Daniel Horn‐Ghetko, David T. Krist, Jesuraj Rajan Prabu et al.*187 citations*
529. **[Baleen whale prey consumption based on high-resolution foraging measurements ↗](https://doi.org/10.1038/s41586-021-03991-5)**Matthew S. Savoca, Max F. Czapanskiy, Shirel R. Kahane‐Rapport et al.*186 citations*
530. **[Five carbon- and nitrogen-bearing species in a hot giant planet’s atmosphere ↗](https://doi.org/10.1038/s41586-021-03381-x)**Paolo Giacobbe, Matteo Brogi, Siddharth Gandhi et al.*186 citations*
531. **[IL-27 signalling promotes adipocyte thermogenesis and energy expenditure ↗](https://doi.org/10.1038/s41586-021-04127-5)**Qian Wang, Dehai Li, Guangchao Cao et al.*186 citations*
532. **[The nutritional quality of cereals varies geospatially in Ethiopia and Malawi ↗](https://doi.org/10.1038/s41586-021-03559-3)**Dawd Gashu, Patson Cleopus Nalivata, Tilahun Amede et al.*186 citations*
533. **[Fc-engineered antibody therapeutics with improved anti-SARS-CoV-2 efficacy ↗](https://doi.org/10.1038/s41586-021-04017-w)**Rachel Yamin, Andrew Jones, Hans-Heinrich Hoffmann et al.*185 citations*
534. **[Cold-induced Arabidopsis FRIGIDA nuclear condensates for FLC repression ↗](https://doi.org/10.1038/s41586-021-04062-5)**Pan Zhu, Clare Lister, Caroline Dean*184 citations*
535. **[Defining genome architecture at base-pair resolution ↗](https://doi.org/10.1038/s41586-021-03639-4)**Hua Peng, Mohsin Badat, Lars L. P. Hanssen et al.*184 citations*
536. **[Inter-cellular CRISPR screens reveal regulators of cancer cell phagocytosis ↗](https://doi.org/10.1038/s41586-021-03879-4)**Roarke A. Kamber, Yoko Nishiga, Bhek Morton et al.*184 citations*
537. **[Structure of Geobacter pili reveals secretory rather than nanowire behaviour ↗](https://doi.org/10.1038/s41586-021-03857-w)**Yangqi Gu, Vishok Srikanth, Aldo I. Salazar-Morales et al.*184 citations*
538. **[Ubiquitous atmospheric production of organic acids mediated by cloud droplets ↗](https://doi.org/10.1038/s41586-021-03462-x)**Bruno Franco, Thomas Blumenstock, Changmin Cho et al.*184 citations*
539. **[A biosensor for the direct visualization of auxin ↗](https://doi.org/10.1038/s41586-021-03425-2)**Ole Herud-Sikimic, André C. Stiel, Martina Kolb et al.*183 citations*
540. **[An engineered IL-2 partial agonist promotes CD8+ T cell stemness ↗](https://doi.org/10.1038/s41586-021-03861-0)**Fei Mo, Zhiya Yu, Peng Li et al.*183 citations*
541. **[NLRs guard metabolism to coordinate pattern- and effector-triggered immunity ↗](https://doi.org/10.1038/s41586-021-04219-2)**Keran Zhai, Di Liang, Helin Li et al.*183 citations*
542. **[Actin cables and comet tails organize mitochondrial networks in mitosis ↗](https://doi.org/10.1038/s41586-021-03309-5)**Andrew S. Moore, Stephen M. Coscia, Cory L. Simpson et al.*182 citations*
543. **[An atlas of gene regulatory elements in adult mouse cerebrum ↗](https://doi.org/10.1038/s41586-021-03604-1)**Yang Eric Li, Sebastian Preißl, Xiaomeng Hou et al.*182 citations*
544. **[A roadmap for the Human Developmental Cell Atlas ↗](https://doi.org/10.1038/s41586-021-03620-1)**Muzlifah A. Haniffa, Deanne M. Taylor, Sten Linnarsson et al.*181 citations*
545. **[Climate policy models need to get real about people — here’s how ↗](https://doi.org/10.1038/d41586-021-01500-2)**Wei Peng, Gokul C. Iyer, Valentina Bosetti et al.*181 citations*
546. **[Pollinators contribute to the maintenance of flowering plant diversity ↗](https://doi.org/10.1038/s41586-021-03890-9)**Na Wei, Rainee L. Kaczorowski, Gerardo Arceo‐Gómez et al.*181 citations*
547. **[A body map of somatic mutagenesis in morphologically normal human tissues ↗](https://doi.org/10.1038/s41586-021-03836-1)**Ruoyan Li, Lin Di, Jie Li et al.*180 citations*
548. **[IPCC climate report: Earth is warmer than it’s been in 125,000 years ↗](https://doi.org/10.1038/d41586-021-02179-1)**Jeff Tollefson*180 citations*
549. **[Non-Hermitian topological whispering gallery ↗](https://doi.org/10.1038/s41586-021-03833-4)**Bolun Hu, Zhiwang Zhang, Haixiao Zhang et al.*180 citations*
550. **[Measurement of gravitational coupling between millimetre-sized masses ↗](https://doi.org/10.1038/s41586-021-03250-7)**Tobias Westphal, Hans Hepach, Jeremias Pfaff et al.*179 citations*
551. **[Control of osteoblast regeneration by a train of Erk activity waves ↗](https://doi.org/10.1038/s41586-020-03085-8)**Alessandro De Simone, Maya N. Evanitsky, Luke Hayden et al.*178 citations*
552. **[Omicron extensively but incompletely escapes Pfizer BNT162b2 neutralization ↗](https://doi.org/10.1038/d41586-021-03824-5)**Sandile Cele, Laurelle Jackson, David S. Khoury et al.*178 citations*
553. **[Peopling of the Americas as inferred from ancient genomics ↗](https://doi.org/10.1038/s41586-021-03499-y)**Eske Willerslev, David J. Meltzer*178 citations*
554. **[Structures of the glucocorticoid-bound adhesion receptor GPR97–Go complex ↗](https://doi.org/10.1038/s41586-020-03083-w)**Yu-Qi Ping, Chunyou Mao, Peng Xiao et al.*177 citations*
555. **[Asymmetric response of interfacial water to applied electric fields ↗](https://doi.org/10.1038/s41586-021-03504-4)**Angelo Montenegro, Chayan Dutta, Muhammet Mammetkuliyev et al.*176 citations*
556. **[COVID’s mental-health toll: how scientists are tracking a surge in depression ↗](https://doi.org/10.1038/d41586-021-00175-z)**Alison Abbott*176 citations*
557. **[Genome surveillance by HUSH-mediated silencing of intronless mobile elements ↗](https://doi.org/10.1038/s41586-021-04228-1)**Marta Seczyńska, Stuart Bloor, Sergio Martínez Cuesta et al.*176 citations*
558. **[Nasal delivery of an IgM offers broad protection from SARS-CoV-2 variants ↗](https://doi.org/10.1038/s41586-021-03673-2)**Zhiqiang Ku, Xuping Xie, Paul R. Hinton et al.*176 citations*
559. **[BARD1 reads H2A lysine 15 ubiquitination to direct homologous recombination ↗](https://doi.org/10.1038/s41586-021-03776-w)**Jordan R. Becker, Gillian Clifford, Clara Bonnet et al.*175 citations*
560. **[Corticosterone inhibits GAS6 to govern hair follicle stem-cell quiescence ↗](https://doi.org/10.1038/s41586-021-03417-2)**Sekyu Choi, Bing Zhang, Sai Ma et al.*175 citations*
561. **[Self-similar mesocrystals form via interface-driven nucleation and assembly ↗](https://doi.org/10.1038/s41586-021-03300-0)**Guomin Zhu, Maria L. Sushko, John S. Loring et al.*175 citations*
562. **[An organoid-based organ-repurposing approach to treat short bowel syndrome ↗](https://doi.org/10.1038/s41586-021-03247-2)**Shinya Sugimoto, Eiji Kobayashi, Masayuki Fujii et al.*174 citations*
563. **[High-fidelity laser-free universal control of trapped ion qubits ↗](https://doi.org/10.1038/s41586-021-03809-4)**R. Srinivas, S. C. Burd, H. M. Knaack et al.*174 citations*
564. **[Mental health concerns during the COVID-19 pandemic as revealed by helpline calls ↗](https://doi.org/10.1038/s41586-021-04099-6)**Marius Brülhart, Valentin Klotzbücher, Rafael Lalive et al.*174 citations*
565. **[Neuroprosthetic baroreflex controls haemodynamics after spinal cord injury ↗](https://doi.org/10.1038/s41586-020-03180-w)**Jordan W. Squair, Matthieu Gautier, Loïs Mahe et al.*174 citations*
566. **[AIM2 in regulatory T cells restrains autoimmune diseases ↗](https://doi.org/10.1038/s41586-021-03231-w)**Wei-Chun Chou, Zengli Guo, Hao Guo et al.*173 citations*
567. **[Tritiation of aryl thianthrenium salts with a molecular palladium catalyst ↗](https://doi.org/10.1038/s41586-021-04007-y)**Da Chuan Zhao, Roland Petzold, Jiyao Yan et al.*173 citations*
568. **[CRL4AMBRA1 is a master regulator of D-type cyclins ↗](https://doi.org/10.1038/s41586-021-03445-y)**Daniele Simoneschi, Gergely Róna, Nan Zhou et al.*172 citations*
569. **[Detection of a particle shower at the Glashow resonance with IceCube ↗](https://doi.org/10.1038/s41586-021-03256-1)**M. G. Aartsen, R. Abbasi, M. Ackermann et al.*172 citations*
570. **[Overwintering fires in boreal forests ↗](https://doi.org/10.1038/s41586-021-03437-y)**Rebecca C. Scholten, Randi R. Jandt, Eric A. Miller et al.*172 citations*
571. **[A phase-separated nuclear GBPL circuit controls immunity in plants ↗](https://doi.org/10.1038/s41586-021-03572-6)**Shuai Huang, Shiwei Zhu, Pradeep Kumar et al.*171 citations*
572. **[Operationalizing the net-negative carbon economy ↗](https://doi.org/10.1038/s41586-021-03723-9)**Johannes Bednar, Michael H. Obersteiner, Artem Baklanov et al.*171 citations*
573. **[Programmable interactions and emergent geometry in an array of atom clouds ↗](https://doi.org/10.1038/s41586-021-04156-0)**Avikar Periwal, Eric S. Cooper, Philipp Kunkel et al.*171 citations*
574. **[Broadly neutralizing antibodies target a haemagglutinin anchor epitope ↗](https://doi.org/10.1038/s41586-021-04356-8)**Jenna J. Guthmiller, Julianna Han, Henry A. Utset et al.*170 citations*
575. **[Direct observation of ultrafast hydrogen bond strengthening in liquid water ↗](https://doi.org/10.1038/s41586-021-03793-9)**Jie Yang, Riccardo Dettori, J. Pedro F. Nunes et al.*170 citations*
576. **[Old-growth forest carbon sinks overestimated ↗](https://doi.org/10.1038/s41586-021-03266-z)**Per Gundersen, Emil Engelund Thybring, Thomas Nord‐Larsen et al.*170 citations*
577. **[A high-resolution picture of kinship practices in an Early Neolithic tomb ↗](https://doi.org/10.1038/s41586-021-04241-4)**Chris Fowler, Íñigo Olalde, Vicki Cummings et al.*169 citations*
578. **[Demonstration of reduced neoclassical energy transport in Wendelstein 7-X ↗](https://doi.org/10.1038/s41586-021-03687-w)**C. D. Beidler, Håkan M. Smith, Arturo Alonso et al.*169 citations*
579. **[Experimental quantum speed-up in reinforcement learning agents ↗](https://doi.org/10.1038/s41586-021-03242-7)**Valeria Saggio, Beate E. Asenbeck, A. Hamann et al.*169 citations*
580. **[How the coronavirus infects cells — and why Delta is so dangerous ↗](https://doi.org/10.1038/d41586-021-02039-y)**Megan Scudellari*169 citations*
581. **[Regulation of prefrontal patterning and connectivity by retinoic acid ↗](https://doi.org/10.1038/s41586-021-03953-x)**Mikihito Shibata, Kartik Pattabiraman, Belén Lorente-Galdós et al.*169 citations*
582. **[The AMBRA1 E3 ligase adaptor regulates the stability of cyclin D ↗](https://doi.org/10.1038/s41586-021-03474-7)**Andrea C. Chaikovsky, Chuan Li, Edwin E. Jeng et al.*167 citations*
583. **[COVID curbed carbon emissions in 2020 — but not by much ↗](https://doi.org/10.1038/d41586-021-00090-3)**Jeff Tollefson*166 citations*
584. **[RNA transcripts stimulate homologous recombination by forming DR-loops ↗](https://doi.org/10.1038/s41586-021-03538-8)**Jian Ouyang, Tribhuwan Yadav, Jiamin Zhang et al.*165 citations*
585. **[Regulation of intestinal immunity and tissue repair by enteric glia ↗](https://doi.org/10.1038/s41586-021-04006-z)**Fränze Progatzky, Michael Shapiro, Song Hui Chng et al.*165 citations*
586. **[A lithium-isotope perspective on the evolution of carbon and silicon cycles ↗](https://doi.org/10.1038/s41586-021-03612-1)**Boriana Kalderon-Asael, Joachim A. R. Katchinoff, Noah J. Planavsky et al.*164 citations*
587. **[Structure of Hsp90–p23–GR reveals the Hsp90 client-remodelling mechanism ↗](https://doi.org/10.1038/s41586-021-04236-1)**Chari M. Noddings, Ray Yu‐Ruei Wang, Jill L. Johnson et al.*164 citations*
588. **[Signatures of moiré trions in WSe2/MoSe2 heterobilayers ↗](https://doi.org/10.1038/s41586-021-03541-z)**Erfu Liu, Elyse Barré, Jeremiah van Baren et al.*163 citations*
589. **[Single-cell Ribo-seq reveals cell cycle-dependent translational pausing ↗](https://doi.org/10.1038/s41586-021-03887-4)**Michael A. VanInsberghe, Jeroen van den Berg, Amanda Andersson-Rolf et al.*163 citations*
590. **[China’s COVID vaccines have been crucial — now immunity is waning ↗](https://doi.org/10.1038/d41586-021-02796-w)**Smriti Mallapaty*161 citations*
591. **[Trapped fractional charges at bulk defects in topological insulators ↗](https://doi.org/10.1038/s41586-020-03117-3)**Christopher W. Peterson, Tianhe Li, Wentao Jiang et al.*161 citations*
592. **[AMBRA1 regulates cyclin D to guard S-phase entry and genomic integrity ↗](https://doi.org/10.1038/s41586-021-03422-5)**Emiliano Maiani, Giacomo Milletti, Francesca Nazio et al.*160 citations*
593. **[Structure of human Cav2.2 channel blocked by the painkiller ziconotide ↗](https://doi.org/10.1038/s41586-021-03699-6)**Shuai Gao, Xia Yin Yao, Nieng Yan*160 citations*
594. **[Subterahertz collective dynamics of polar vortices ↗](https://doi.org/10.1038/s41586-021-03342-4)**Qian Li, Vladimir A. Stoica, Marek Paściak et al.*160 citations*
595. **[Blood and immune development in human fetal bone marrow and Down syndrome ↗](https://doi.org/10.1038/s41586-021-03929-x)**Laura Jardine, Simone Webb, Issac Goh et al.*159 citations*
596. **[Phosphorus-mediated sp2–sp3 couplings for C–H fluoroalkylation of azines ↗](https://doi.org/10.1038/s41586-021-03567-3)**Xuan Zhang, Kyle G. Nottingham, Chirag Patel et al.*159 citations*
597. **[Platypus and echidna genomes reveal mammalian biology and evolution ↗](https://doi.org/10.1038/s41586-020-03039-0)**Yang Zhou, Linda Shearwin‐Whyatt, Jing Li et al.*159 citations*
598. **[Structure, function and pharmacology of human itch receptor complexes ↗](https://doi.org/10.1038/s41586-021-04077-y)**Fan Yang, Lulu Guo, Yu Li et al.*159 citations*
599. **[Experimental observation of non-Abelian topological charges and edge states ↗](https://doi.org/10.1038/s41586-021-03521-3)**Qinghua Guo, Tianshu Jiang, Ruo-Yang Zhang et al.*158 citations*
600. **[Lineage tracing of human development through somatic mutations ↗](https://doi.org/10.1038/s41586-021-03548-6)**Michael Spencer Chapman, Anna Maria Ranzoni, Brynelle Myers et al.*158 citations*

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