---
title: "Proceedings of the National Academy of Sciences 2020 — all"
description: "All 4,216 papers of Proceedings of the National Academy of Sciences 2020 · page 1 of 43, the most cited first; those with our summary link to it."
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# Proceedings of the National Academy of Sciences 2020: all accepted papers

4,216 papers · page 1 of 43 · the most cited first · [← Proceedings of the National Academy of Sciences 2020](https://md.research.elamur.ai/free/en/venues/proceedings-of-the-national-academy-of-sciences/2020/)

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1. **[RepeatModeler2 for automated genomic discovery of transposable element families ↗](https://doi.org/10.1073/pnas.1921046117)**Jullien M. Flynn, Robert M. Hubley, Clément Goubert et al.*3,800 citations*
2. **[Cell entry mechanisms of SARS-CoV-2 ↗](https://doi.org/10.1073/pnas.2003138117)**Jian Shang, Yushun Wan, Chuming Luo et al.*3,130 citations*
3. **[Effective treatment of severe COVID-19 patients with tocilizumab ↗](https://doi.org/10.1073/pnas.2005615117)**Xiaoling Xu, Mingfeng Han, Tiantian Li et al.*2,647 citations*
4. **[Effectiveness of convalescent plasma therapy in severe COVID-19 patients ↗](https://doi.org/10.1073/pnas.2004168117)**Kai Duan, Bende Liu, Cesheng Li et al.*1,960 citations*
5. **[The impact of COVID-19 on small business outcomes and expectations ↗](https://doi.org/10.1073/pnas.2006991117)**Alexander Wickman Bartik, Marianne Bertrand, Zoë Cullen et al.*1,604 citations*
6. **[Land use and climate change impacts on global soil erosion by water (2015-2070) ↗](https://doi.org/10.1073/pnas.2001403117)**Pasquale Borrelli, David A. Robinson, Panos Panagos et al.*1,465 citations*
7. **[Enhanced isolation of SARS-CoV-2 by TMPRSS2-expressing cells ↗](https://doi.org/10.1073/pnas.2002589117)**Shutoku Matsuyama, Naganori Nao, Kazuya Shirato et al.*1,411 citations*
8. **[Improved protein structure prediction using predicted interresidue orientations ↗](https://doi.org/10.1073/pnas.1914677117)**Jianyi Yang, Ivan V. Anishchenko, Hahnbeom Park et al.*1,411 citations*
9. **[Active learning narrows achievement gaps for underrepresented students in undergraduate science, technology, engineering, and math ↗](https://doi.org/10.1073/pnas.1916903117)**Elli J. Theobald, Mariah J. Hill, Elisa T. Tran et al.*1,366 citations*
10. **[Identifying airborne transmission as the dominant route for the spread of COVID-19 ↗](https://doi.org/10.1073/pnas.2009637117)**Renyi Zhang, Yixin Li, Annie Li Zhang et al.*1,235 citations*
11. **[COVID-19 lockdowns cause global air pollution declines ↗](https://doi.org/10.1073/pnas.2006853117)**Zander Samuel Venter, Kristin Aunan, Sourangsu Chowdhury et al.*1,081 citations*
12. **[The airborne lifetime of small speech droplets and their potential importance in SARS-CoV-2 transmission ↗](https://doi.org/10.1073/pnas.2006874117)**Valentyn Stadnytskyi, Christina E. Bax, Ad Bax et al.*1,077 citations*
13. **[Spread and dynamics of the COVID-19 epidemic in Italy: Effects of emergency containment measures ↗](https://doi.org/10.1073/pnas.2004978117)**Marino Gatto, Enrico Bertuzzo, Lorenzo Mari et al.*1,051 citations*
14. **[Syrian hamsters as a small animal model for SARS-CoV-2 infection and countermeasure development ↗](https://doi.org/10.1073/pnas.2009799117)**Masaki Imai, Kiyoko Iwatsuki‐Horimoto, Masato Hatta et al.*1,021 citations*
15. **[Phylogenetic network analysis of SARS-CoV-2 genomes ↗](https://doi.org/10.1073/pnas.2004999117)**Peter Forster, Lucy Forster, Colin Renfrew et al.*1,020 citations*
16. **[The frontier of simulation-based inference ↗](https://doi.org/10.1073/pnas.1912789117)**Kyle Stuart Cranmer, Johann Brehmer, Gilles Louppe*1,020 citations*
17. **[Economic and social consequences of human mobility restrictions under COVID-19 ↗](https://doi.org/10.1073/pnas.2007658117)**Giovanni Bonaccorsi, Francesco Pierri, Matteo Cinelli et al.*1,002 citations*
18. **[The Diversity–Innovation Paradox in Science ↗](https://doi.org/10.1073/pnas.1915378117)**Bas Hofstra, Vivek Kulkarni, Sebastian Munoz-Najar Galvez et al.*975 citations*
19. **[Oncogenic activation of PI3K-AKT-mTOR signaling suppresses ferroptosis via SREBP-mediated lipogenesis ↗](https://doi.org/10.1073/pnas.2017152117)**Jun-Mei Yi, Jiajun Zhu, Jiao Wu et al.*974 citations*
20. **[Historical comparison of gender inequality in scientific careers across countries and disciplines ↗](https://doi.org/10.1073/pnas.1914221117)**Junming Huang, Alexander J. Gates, Roberta Sinatra et al.*954 citations*
21. **[Social tipping dynamics for stabilizing Earth’s climate by 2050 ↗](https://doi.org/10.1073/pnas.1900577117)**Ilona M. Otto, Jonathan F. Donges, Roger Cremades et al.*954 citations*
22. **[Vertebrates on the brink as indicators of biological annihilation and the sixth mass extinction ↗](https://doi.org/10.1073/pnas.1922686117)**Gerardo Ceballos, Paul R. Ehrlich, Peter H. Raven*918 citations*
23. **[Demographic science aids in understanding the spread and fatality rates of COVID-19 ↗](https://doi.org/10.1073/pnas.2004911117)**Jennifer B. Dowd, Liliana Andriano, David M. Brazel et al.*905 citations*
24. **[Prophylactic and therapeutic remdesivir (GS-5734) treatment in the rhesus macaque model of MERS-CoV infection ↗](https://doi.org/10.1073/pnas.1922083117)**Emmie de Wit, Friederike Feldmann, Jacqueline Cronin et al.*887 citations*
25. **[Gender differences in COVID-19 attitudes and behavior: Panel evidence from eight countries ↗](https://doi.org/10.1073/pnas.2012520117)**Vincenzo Galasso, Vincent Pons, Paola Antonia Profeta et al.*867 citations*
26. **[Future of the human climate niche ↗](https://doi.org/10.1073/pnas.1910114117)**Chi Xu, Timothy A. Kohler, Timothy M. Lenton et al.*852 citations*
27. **[Effective fisheries management instrumental in improving fish stock status ↗](https://doi.org/10.1073/pnas.1909726116)**Ray W. Hilborn, Ricardo O. Amoroso, Christopher M. Anderson et al.*850 citations*
28. **[RCP8.5 tracks cumulative CO 2 emissions ↗](https://doi.org/10.1073/pnas.2007117117)**Christopher R. Schwalm, Spencer Glendon, Philip B. Duffy*804 citations*
29. **[Large stocks of peatland carbon and nitrogen are vulnerable to permafrost thaw ↗](https://doi.org/10.1073/pnas.1916387117)**Gustaf Hugelius, Julie Loisel, Sarah Chadburn et al.*784 citations*
30. **[On instabilities of deep learning in image reconstruction and the potential costs of AI ↗](https://doi.org/10.1073/pnas.1907377117)**Vegard Antun, Francesco Renna, Clarice Poon et al.*779 citations*
31. **[A digital media literacy intervention increases discernment between mainstream and false news in the United States and India ↗](https://doi.org/10.1073/pnas.1920498117)**Andrew M. Guess, Michael Lerner, Benjamin A. Lyons et al.*762 citations*
32. **[Racial disparities in automated speech recognition ↗](https://doi.org/10.1073/pnas.1915768117)**Allison Koenecke, Andrew Nam, Emily Lake et al.*754 citations*
33. **[Recent responses to climate change reveal the drivers of species extinction and survival ↗](https://doi.org/10.1073/pnas.1913007117)**Cristian Román‐Palacios, John J. Wiens*744 citations*
34. **[Physician–patient racial concordance and disparities in birthing mortality for newborns ↗](https://doi.org/10.1073/pnas.1913405117)**Brad N. Greenwood, Rachel R. Hardeman, Laura Huang et al.*740 citations*
35. **[The process of Lewy body formation, rather than simply α-synuclein fibrillization, is one of the major drivers of neurodegeneration ↗](https://doi.org/10.1073/pnas.1913904117)**Anne‐Laure Mahul‐Mellier, Johannes Burtscher, Niran Maharjan et al.*700 citations*
36. **[Gender imbalance in medical imaging datasets produces biased classifiers for computer-aided diagnosis ↗](https://doi.org/10.1073/pnas.1919012117)**Agostina J. Larrazabal, Nicolás Nieto, Victoria Peterson et al.*680 citations*
37. **[Microbial tryptophan metabolites regulate gut barrier function via the aryl hydrocarbon receptor ↗](https://doi.org/10.1073/pnas.2000047117)**Samantha A. Scott, Jingjing Fu, Pamela V. Chang*673 citations*
38. **[Global increase in major tropical cyclone exceedance probability over the past four decades ↗](https://doi.org/10.1073/pnas.1920849117)**James P. Kossin, Kenneth R. Knapp, Timothy L. Olander et al.*664 citations*
39. **[Cell profiling of mouse acute kidney injury reveals conserved cellular responses to injury ↗](https://doi.org/10.1073/pnas.2005477117)**Yuhei Kirita, Hao Wu, Kohei Uchimura et al.*641 citations*
40. **[Broad host range of SARS-CoV-2 predicted by comparative and structural analysis of ACE2 in vertebrates ↗](https://doi.org/10.1073/pnas.2010146117)**Joana Damas, Graham M. Hughes, Kathleen C. Keough et al.*630 citations*
41. **[Rhizosphere microbiome mediates systemic root metabolite exudation by root-to-root signaling ↗](https://doi.org/10.1073/pnas.1912130117)**Elisa Korenblum, Yonghui Dong, Jędrzej Szymański et al.*597 citations*
42. **[The impact of COVID-19 nonpharmaceutical interventions on the future dynamics of endemic infections ↗](https://doi.org/10.1073/pnas.2013182117)**Rachel E. Baker, Sang Woo Park, Wenchang Yang et al.*587 citations*
43. **[ALKBH5 regulates anti–PD-1 therapy response by modulating lactate and suppressive immune cell accumulation in tumor microenvironment ↗](https://doi.org/10.1073/pnas.1918986117)**Na Li, Yuqi Kang, Lingling Wang et al.*576 citations*
44. **[The challenges of modeling and forecasting the spread of COVID-19 ↗](https://doi.org/10.1073/pnas.2006520117)**Andrea Louise Bertozzi, Elisa Franco, George Mohler et al.*562 citations*
45. **[Observation of non-Hermitian topology and its bulk–edge correspondence in an active mechanical metamaterial ↗](https://doi.org/10.1073/pnas.2010580117)**Ananya Ghatak, Martin Brandenbourger, Jasper van Wezel et al.*558 citations*
46. **[Impact of international travel and border control measures on the global spread of the novel 2019 coronavirus outbreak ↗](https://doi.org/10.1073/pnas.2002616117)**Chad R. Wells, Pratha Sah, Seyed M. Moghadas et al.*553 citations*
47. **[Biologically inspired flexible photonic films for efficient passive radiative cooling ↗](https://doi.org/10.1073/pnas.2001802117)**Haiwen Zhang, Kally Chein Sheng Ly, Xianghui Liu et al.*550 citations*
48. **[Benign overfitting in linear regression ↗](https://doi.org/10.1073/pnas.1907378117)**Peter L. Bartlett, Philip M. Long, Gábor Lugosi et al.*547 citations*
49. **[Using gross ecosystem product (GEP) to value nature in decision making ↗](https://doi.org/10.1073/pnas.1911439117)**Zhiyun Ouyang, Changsu Song, Hua Zheng et al.*543 citations*
50. **[The European Union Emissions Trading System reduced CO 2 emissions despite low prices ↗](https://doi.org/10.1073/pnas.1918128117)**Patrick J. Bayer, Michaël Aklin*538 citations*
51. **[Political Partisanship Influences Behavioral Responses to Governors’ Recommendations for COVID-19 Prevention in the United States ↗](https://doi.org/10.1073/pnas.2007835117)**Guy Grossman, Soojong Kim, Jonah M. Rexer et al.*532 citations*
52. **[SARS-CoV-2 Orf6 hijacks Nup98 to block STAT nuclear import and antagonize interferon signaling ↗](https://doi.org/10.1073/pnas.2016650117)**Lisa Miorin, Thomas Kehrer, María Teresa Sánchez-Aparicio et al.*527 citations*
53. **[Characterization and engineering of a two-enzyme system for plastics depolymerization ↗](https://doi.org/10.1073/pnas.2006753117)**Brandon C. Knott, Erika Erickson, Mark D. Allen et al.*526 citations*
54. **[Impacts of current and future large dams on the geographic range connectivity of freshwater fish worldwide ↗](https://doi.org/10.1073/pnas.1912776117)**Valerio Barbarossa, Rafael Schmitt, Mark A. J. Huijbregts et al.*496 citations*
55. **[Projecting hospital utilization during the COVID-19 outbreaks in the United States ↗](https://doi.org/10.1073/pnas.2004064117)**Seyed M. Moghadas, Affan Shoukat, Meagan C. Fitzpatrick et al.*494 citations*
56. **[Critical period regulation across multiple timescales ↗](https://doi.org/10.1073/pnas.1820836117)**Rebecca K. Reh, Brian George Dias, Charles A. Nelson et al.*487 citations*
57. **[Polyunsaturated fatty acid biosynthesis pathway determines ferroptosis sensitivity in gastric cancer ↗](https://doi.org/10.1073/pnas.2006828117)**Ji-Yoon Lee, Miso Nam, Hye Young Son et al.*477 citations*
58. **[Progress toward gender equality in the United States has slowed or stalled ↗](https://doi.org/10.1073/pnas.1918891117)**Paula England, Andrew Levine, Emma Mishel*475 citations*
59. **[Single-nucleus transcriptome analysis reveals dysregulation of angiogenic endothelial cells and neuroprotective glia in Alzheimer’s disease ↗](https://doi.org/10.1073/pnas.2008762117)**Shun‐Fat Lau, Han Cao, Amy K.Y. Fu et al.*471 citations*
60. **[A scalable pipeline for designing reconfigurable organisms ↗](https://doi.org/10.1073/pnas.1910837117)**Sam Kriegman, Douglas J. Blackiston, Michael G. Levin et al.*461 citations*
61. **[TLR signaling adapter BCAP regulates inflammatory to reparatory macrophage transition by promoting histone lactylation ↗](https://doi.org/10.1073/pnas.2009778117)**Ricardo A. Irizarry-Caro, Margaret M. McDaniel, Garrett R. Overcast et al.*457 citations*
62. **[The implications of silent transmission for the control of COVID-19 outbreaks ↗](https://doi.org/10.1073/pnas.2008373117)**Seyed M. Moghadas, Meagan C. Fitzpatrick, Pratha Sah et al.*449 citations*
63. **[The functional activity of E-cadherin controls tumor cell metastasis at multiple steps ↗](https://doi.org/10.1073/pnas.1918167117)**Tae-Young Na, Leslayann C. Schecterson, Alisha M. Mendonsa et al.*441 citations*
64. **[Predicting long-term dynamics of soil salinity and sodicity on a global scale ↗](https://doi.org/10.1073/pnas.2013771117)**Amirhossein Hassani, Adisa Azapagic, Nima Shokri*440 citations*
65. **[BCG vaccine protection from severe coronavirus disease 2019 (COVID-19) ↗](https://doi.org/10.1073/pnas.2008410117)**Luis E. Escobar, Alvaro Molina-Cruz, Carolina V. Barillas-Mury*437 citations*
66. **[First-principles experimental demonstration of ferroelectricity in a thermotropic nematic liquid crystal: Polar domains and striking electro-optics ↗](https://doi.org/10.1073/pnas.2002290117)**Xi Chen, Eva D. Korblova, Dengpan Dong et al.*436 citations*
67. **[NETosis proceeds by cytoskeleton and endomembrane disassembly and PAD4-mediated chromatin decondensation and nuclear envelope rupture ↗](https://doi.org/10.1073/pnas.1909546117)**Hawa Racine Thiam, Siu Ling Wong, Rong Qiu et al.*433 citations*
68. **[Land-use intensity alters networks between biodiversity, ecosystem functions, and services ↗](https://doi.org/10.1073/pnas.2016210117)**María R. Felipe‐Lucia, Santiago Soliveres, Caterina Penone et al.*432 citations*
69. **[Experimental infection of domestic dogs and cats with SARS-CoV-2: Pathogenesis, transmission, and response to reexposure in cats ↗](https://doi.org/10.1073/pnas.2013102117)**Angela M. Bosco‐Lauth, Airn E. Hartwig, Stephanie M. Porter et al.*424 citations*
70. **[Rapid isothermal amplification and portable detection system for SARS-CoV-2 ↗](https://doi.org/10.1073/pnas.2014739117)**Anurup Ganguli, Ariana Mostafa, Jacob Berger et al.*421 citations*
71. **[Global diversity of microbial communities in marine sediment ↗](https://doi.org/10.1073/pnas.1919139117)**Tatsuhiko Hoshino, Hideyuki Doi, Go‐Ichiro Uramoto et al.*412 citations*
72. **[Face masks considerably reduce COVID-19 cases in Germany ↗](https://doi.org/10.1073/pnas.2015954117)**Timo Mitze, Reinhold Kosfeld, Johannes Rode et al.*411 citations*
73. **[Systemic signaling during abiotic stress combination in plants ↗](https://doi.org/10.1073/pnas.2005077117)**Sara I. Zandalinas, Yosef Fichman, Amith R. Devireddy et al.*410 citations*
74. **[Social distancing responses to COVID-19 emergency declarations strongly differentiated by income ↗](https://doi.org/10.1073/pnas.2009412117)**Joakim Weill, Matthieu Stigler, Olivier Deschênes et al.*406 citations*
75. **[Worldwide carrier frequency and genetic prevalence of autosomal recessive inherited retinal diseases ↗](https://doi.org/10.1073/pnas.1913179117)**Mor Hanany, Carlo Rivolta, Dror Sharon*401 citations*
76. **[Spatially separating redox centers on 2D carbon nitride with cobalt single atom for photocatalytic H 2 O 2 production ↗](https://doi.org/10.1073/pnas.1913403117)**Chiheng Chu, Qianhong Zhu, Zhenhua Pan et al.*400 citations*
77. **[How cholesterol stiffens unsaturated lipid membranes ↗](https://doi.org/10.1073/pnas.2004807117)**Saptarshi Chakraborty, Milka Doktorova, Trivikram R. Molugu et al.*397 citations*
78. **[Superhydrophobic photothermal icephobic surfaces based on candle soot ↗](https://doi.org/10.1073/pnas.2001972117)**Shuwang Wu, Yingjie Du, Yousif Alsaid et al.*395 citations*
79. **[Systemic complement activation is associated with respiratory failure in COVID-19 hospitalized patients ↗](https://doi.org/10.1073/pnas.2010540117)**Jan Cato Holter, Søren Erik Pischke, Eline de Boer et al.*395 citations*
80. **[Prevalence of neural collapse during the terminal phase of deep learning training ↗](https://doi.org/10.1073/pnas.2015509117)**Vardan Papyan, X. Y. Han, David L. Donoho*393 citations*
81. **[Superantigenic character of an insert unique to SARS-CoV-2 spike supported by skewed TCR repertoire in patients with hyperinflammation ↗](https://doi.org/10.1073/pnas.2010722117)**Mary Hongying Cheng, She Zhang, Rebecca A. Porritt et al.*393 citations*
82. **[Impacts of the COVID-19 pandemic on rural America ↗](https://doi.org/10.1073/pnas.2019378118)**J. Tom Mueller, Kathryn McConnell, Paul Berne Burow et al.*391 citations*
83. **[Instant tough bioadhesive with triggerable benign detachment ↗](https://doi.org/10.1073/pnas.2006389117)**Xiaoyu Chen, Hyunwoo Yuk, Jingjing Wu et al.*391 citations*
84. **[Viral zoonotic risk is homogenous among taxonomic orders of mammalian and avian reservoir hosts ↗](https://doi.org/10.1073/pnas.1919176117)**Nardus Mollentze, Daniel G. Streicker*384 citations*
85. **[COVID-19 lockdown induces disease-mitigating structural changes in mobility networks ↗](https://doi.org/10.1073/pnas.2012326117)**Frank Schlosser, Benjamin F. Maier, Olivia Jack et al.*379 citations*
86. **[A new malaria vector in Africa: Predicting the expansion range of Anopheles stephensi and identifying the urban populations at risk ↗](https://doi.org/10.1073/pnas.2003976117)**Marianne E Sinka, Samuel Pironon, N. Claire Massey et al.*376 citations*
87. **[TDP-43 α-helical structure tunes liquid–liquid phase separation and function ↗](https://doi.org/10.1073/pnas.1912055117)**Alexander E. Conicella, Gregory L. Dignon, Gül H. Zerze et al.*375 citations*
88. **[In situ sprayed NIR-responsive, analgesic black phosphorus-based gel for diabetic ulcer treatment ↗](https://doi.org/10.1073/pnas.2016268117)**Jiang Ouyang, Xiaoyuan Ji, Xingcai Zhang et al.*371 citations*
89. **[Bioinspired underwater locomotion of light-driven liquid crystal gels ↗](https://doi.org/10.1073/pnas.1917952117)**Hamed Shahsavan, Amirreza Aghakhani, Hao Zeng et al.*370 citations*
90. **[Overstated carbon emission reductions from voluntary REDD+ projects in the Brazilian Amazon ↗](https://doi.org/10.1073/pnas.2004334117)**Thales A. P. West, Jan Börner, Erin O. Sills et al.*368 citations*
91. **[Widespread occurrence of the droplet state of proteins in the human proteome ↗](https://doi.org/10.1073/pnas.2007670117)**Maarten C. Hardenberg, Attila Horváth, Viktor Ambrus et al.*368 citations*
92. **[Understanding the role of individual units in a deep neural network ↗](https://doi.org/10.1073/pnas.1907375117)**David Anthony Bau, Jun-Yan Zhu, Hendrik Strobelt et al.*367 citations*
93. **[The carbon footprint of household energy use in the United States ↗](https://doi.org/10.1073/pnas.1922205117)**Benjamin P. Goldstein, Dimitrios Gounaridis, Joshua P. Newell*363 citations*
94. **[Noninvasive hippocampal blood−brain barrier opening in Alzheimer’s disease with focused ultrasound ↗](https://doi.org/10.1073/pnas.2002571117)**Ali R. Rezai, Manish Ranjan, Pierre-François D’Haese et al.*361 citations*
95. **[Tracking the reach of COVID-19 kin loss with a bereavement multiplier applied to the United States ↗](https://doi.org/10.1073/pnas.2007476117)**Ashton M. Verdery, Emily Smith‐Greenaway, Rachel Margolis et al.*359 citations*
96. **[Molecular adaptations of the blood–brain barrier promote stress resilience vs. depression ↗](https://doi.org/10.1073/pnas.1914655117)**Katarzyna Dudek, Laurence Dion‐Albert, Manon Lebel et al.*357 citations*
97. **[Most invasive species largely conserve their climatic niche ↗](https://doi.org/10.1073/pnas.2004289117)**Chunlong Liu, Christian Wolter, Weiwei Xian et al.*354 citations*
98. **[Extraction of mechanical properties of materials through deep learning from instrumented indentation ↗](https://doi.org/10.1073/pnas.1922210117)**Lu Lu, Ming Dao, Punit Kumar et al.*352 citations*
99. **[Split-TurboID enables contact-dependent proximity labeling in cells ↗](https://doi.org/10.1073/pnas.1919528117)**Kelvin Frank Cho, Tess Caroline Branon, Sanjana Rajeev et al.*352 citations*
100. **[The unreasonable effectiveness of deep learning in artificial intelligence ↗](https://doi.org/10.1073/pnas.1907373117)**Terrence J. Sejnowski*350 citations*

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